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PA15 and Dealing with estate

My Aunt passed away in July leaving a Will the executor named in the Will no longer wishes to carry out the duties. They have filled in a PA15 form can the 4 beneficiary's  of the Will myself being one go head and start to deal with the estate or do we need to fill in a form to get permission to do so. As i understand it the PA15 needs to submitted with probate or does this need to be sent before?
Inheritance tax has me slightly confused is it £325,000 or £500,000? 
A Property in Portugal is involved so guessing this will make it a more complicated process as I image they have different rules than the UK.

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